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Don't know if it has crash protection, hasn't happened with either program for me yet. Desktop runs like Photoshop, save before you close. Autosave - Store always autosaves the canvas whenever it is closed and resumes on open.Layers Window - It is absolute shit on the Desktop version when trying to use the pen since it doesn't distinguish between finger and pen so you have to use the tiny scroll bar which can be difficult.Especially neat are the menus you can make entirely to your own liking with the side button on the surface pen (right click). Desktop has some customization, hotkey reprogramming and Lagoon customizing. Customization/Settings/Hotkeys - Pretty much ZERO settings and options in Store version, kind of odd.My biggest frustration when pinch zooming. Finger Drawing - Can't seem to disable in Desktop like you can in Store.Desktop is adjustable around 200 colours visible, 180 with HSL sliders open. Colour Palette Window - Store version is non-adjustable in length, max 20 visible palette colours.Desktop also has the 'lagoon' menu that sits in a corner but I don't find a need for it. Store has additional colour adjustment: hue/saturation/lightness and colour balance (shadows/mids/highlights). Tools - Desktop has an additional non-automatic 'steady stroke', french curve & adjustable perspective lines.Also you can make accidental brush strokes very easily on Desktop. 2-Finger Zooming - A lot smoother on the Store.
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UI - Store is more rounded and tablet app feeling, as Desktop is more traditional windows.I've mostly stuck with portrait digital painting for now, this was done on Sketchbook Desktop Pro. I've used both for a few months now, first with the Store version and then with the Desktop version.